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Arrange a rack in the middle of the oven and heat to 400°F.Heat 1 tablespoon of the oil in a large skillet over medium heat until shimmering. Add the onions and cook until the onions are sweated and are just starting to turn translucent, about 5 minutes. Add the mushrooms and cook until the mushrooms have released their liquid and most of that liquid has evaporated, 5 to 8 minutes. Add the eggplant, zucchini, and bell pepper, and cook until all are softened, 3 to 5 minutes more.Push the vegetables aside to clear out some space in the middle of the pan. Add 1 teaspoon of the oil, garlic, salt, thyme, oregano, and red pepper flakes if using. Stir the garlic until it is fragrant and just starting to turn golden, about 30 seconds, then stir it into the vegetables. Add the spinach and stir until wilted, 3 to 5 minutes. Remove from the heat.Spread a thin layer of the marinara sauce in the bottom of a 9x13-inch baking dish. Lay 5 of the lasagna noodles over the sauce, breaking them if needed to create a single layer. Spread 1 cup of the ricotta over the noodles. Top with 1/2 of the vegetables, 1 cup of the marinara sauce, and 1 cup of the mozzarella.Arrange another 5 noodles over mozzarella, followed 1 cup of ricotta, the remaining vegetables, 1 cup of the marinara, and 1 cup of the mozzarella. Top this with a final layer of 5 noodles and the remaining sauce. (Reserve the remaining mozzarella for the end of cooking.) Cover the dish tightly with aluminum foil.Bake for 1 hour. Check to make sure the noodles are done by poking the lasagna with a knife; the knife should slide easily through all the layers. If it doesn't, cover and cook for 15 minutes more. When the noodles are done, sprinkle the remaining 1 cup of mozzarella over the top. Bake, uncovered, until the mozzarella has melted, 10 to 15 minutes more.Turn on the broiler and arrange an oven rack 6 inches below the heating element. Broil the lasagna until the mozzarella is browned and bubbling, 1 to 2 minutes.Remove the lasagna from the oven and let it cool on a wire rack for at least 15 minutes before serving.
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