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Preheat oven to 425F. Line a muffin pan with paper liners and set aside.
In a medium bowl, toss together the flour, poppy seeds, baking powder, baking soda, and salt. Set aside.
In a large bowl, combine the sugar and lemon zest. Rub the mixture between your fingers to release the oil. The sugar will be pale yellow and very fragrant.
Then whisk in the melted butter, eggs, sour cream, fresh lemon juice, milk, and vanilla extract.
Add the dry ingredients to the wet ingredients and mix together until just combined. Spoon the batter into the prepared muffin pan.
Bake at 425F for 5 minutes, then reduce the oven to 350F and bake for another 15 minutes or until a toothpick inserted into the center comes out clean.