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Step 1
Position a rack in the center of the oven and preheat to 350ºF.
Step 2
Line an 8-inch square pan with parchment paper on all sides.
Step 3
In a large, deep bowl, rub the lemon zest into the sugar until the sugar feels damp.
Step 4
Whisk in the eggs to combine, then whisk in the oil, sour cream, and lemon juice until smooth.
Step 5
Place a strainer over the bowl and sift in the sweet rice, millet, oat, and tapioca flours with the baking powder, baking soda, and salt.
Step 6
Whisk until combined, then whisk in the poppy seeds.
Step 7
Pour the batter into the prepared pan and spread smooth.
Step 8
Bake until a toothpick inserted in the center comes out clean or with a few moist crumbs, 30-35 minutes.
Step 9
Let the cake cool to room temperature, 1-2 hours. Turn the cake out, peel away the parchment, and place the cake upright on a serving platter or board.
Step 10
Spread the cake with the frosting, decorate the top with poppy seeds, lemon zest, or lemon slices, and serve!